Character list and data matrix:

 [1] Premaxillary teeth present (0), absent (1) (irreversible)
 [2] Parapophyses located in the cranial (0) or central (1) part of the bodies of dorsal vertebrae
 [3] Fibula does (0) not (1) reach calcaneum
 [4] Humeral head cranially concave and caudally convex (0) or globose (1)
 [5] Scapula and coracoid sutured or fused (0), forming a flat joint (1), a joint coracoid in scapula (2), a shallow joint scapula in coracoid (3) or a deep joint scapula in coracoid (4) (see stepmatrix)
 [6] Supracoracoid nerve foramen not (0) opening into an elongate furrow medially and separated from the medial margin of the coracoid by a thick, bony bar (1)
 [7] Broad, deep fossa on the dorsal side of the coracoid: absent (0), present (1)
 [8] Supracoracoid nerve foramen opens above the coracoidal fossa (0) or inside it (1)
 [9] Prominent scapular acromion (0) absent, costolaterally not (1) wider than deep (2) (ordered)
[10] Costal surface of scapular blade not (0) with a prominent, longitudinal furrow (1)
[11] Dorsal and ventral margins of the furcula: subequal in width (0), ventral margin clearly wider than the dorsal margin (1)
[12] Hypocleideum: absent or poorly developed (0) or well developed (1)
[13] Ossified sternal keel: absent (0), not (1) reaching the cranial border of the sternum or beyond it (2) (ordered)
[14] Manual digit I reaches beyond mc II (0), +- reaches its end (1) or is markedly shorter (2) (ordered)
[15] Manual digit III: 4 separate phalanges (0), 3 (1), 2 (2), 1 (3) (irreversible)
[16] Coracoid without (0), with a tiny (1) or with a cylindrical part longer than 1/3 of the coracoid (2) (ordered)
[17] Dorsal margin of the humeral head not (0) concave in its central portion, rising ventrally and dorsally (1)
[18] Prominent bicipital crest of humerus, cranioventrally projecting: absent (0), present (1)
[19] Ventral face of the humeral bicipital crest not (0) with a small fossa for muscular attachment (1)
[20] Distal end of humerus not (0) very compressed craniocaudally (1)
[21] Dorsal cotyla of ulna not (0) separated from the olecranon by a groove (1)
[22] Shaft of radius not (0) with a long longitudinal groove on its caudoventral surface (1)
[23] Pygostyle: absent (0), middle (1), long (2), short (3) (see stepmatrix)
[24] Mc III not (0) projecting distally more than mc II (1)
[25] Manual digit II: proximal phalanx shorter (0) or longer (1) than intermediate phalanx
[26] Pubic foot: present (0), absent (1)
[27] Femoral posterior trochanter: absent or weakly developed (0), hypertrophied (1)
[28] Caudal projection of the lateral border of the distal end of the femur: absent (0), present (1)
[29] Very narrow, deep intercondylar sulcus on tibiotarsus that proximally undercuts condyles: absent (0), present (1)
[30] Trochlea of mt II not (0) much broader than the trochlea of mt III (1)
[31] Mt IV not (0) significantly thinner than mt II & III (1)
[32] Medial rim of the trochlea of mt III with a strong plantar projection: absent (0), present (1)
[33] Mt I not (0) distinctly J-shaped in medial aspect (1)
[34] Mt V present (0), absent (1) (irreversible)
[35] Strong transverse convexity of the dorsal surface of the midshaft of mt III: absent (0), present (1)
[36] Distal end of mt II not (0) strongly curved medially (1)
[37] Sacral vertebrae: 5 (0), 6 (1), 7 (2), 8 (3), 9 (4), 10 or more (5) (ordered)
[38] Pedal digit II without the claw not (0) significantly shorter than IV (1)
[39] Largest pedal claw on digit III (0), II (1), I (2) (unordered)
[40] Pedal digit II not (0) hyperextensible (1)
[41] Distal fusion between mc II and III: absent (0), present (1)
[42] Distal fusion between mt II, III and IV: absent (0), ontogenetically after (1) or before proximal fusion (2) (unordered)
[43] Patella(r groove on femur): absent (0), present (1)
[44] Hypotarsus: absent (0), present (1)
[45] Proximal end of mt III not (0) broadened (1), displaced plantarly (2) or narrowed (subarctometatarsal condition) (3) (see stepmatrix)
[46] Middle tail vertebrae not (0) at least twice as long as the proximal ones (1)
[47] Extensor process on mc I: absent (0), present (1)
[48] Sterna sutured (0) or fused (1)
[49] Procoracoid process: absent (0), present (1)
[50] Pubic symphysis more (0), less than 1/3 of pubic length (1) or absent (2) (irreversible)
[51] Femoral neck: present (0), absent (1)
[52] More (0) or fewer (1) than 10 free caudal vertebrae (irreversible)
[53] Preacetabular process of ilium less than (0) twice as long as postacetabular process (1)
[54] Bicipital scar on ulna not (0) developed as a tubercle (1)
[55] Ventral tubercle of humerus: absent (0), present (1)
[56] Coracoid less (0), more (1) than 1.75 times as long (glenoid to sternum) as broad (maximal distance craniomedial to caudolateral margin)
[57] Articular pneumaticity: absent (0), present (1)
[58] Intercotylar eminence: absent (0), present (1)
[59] Fossa for mt I on distal plantar surface of mt II: absent (0), present (1)
[60] Flange on phalanx II-1 (for attachment of remiges): absent (0), present (1)
[61] Ilia: close together over entire length (0), far apart and vertical over entire length (1) or sigmoid, close together cranially to the acetabulum and far apart caudally (2) (unordered)
[62] Pubic peduncle of ilium about 1 (0) or 3 (1) times as long craniocaudally as the ischiadic one (measured at the base = dorsal end)
[63] Transition point near the middle (1), base (0) or end of the tail (2) (ordered)
[64] Tail distal to transition point has: short, broad prz (or none) (0), short, narrow prz (1), long prz and poz (2) (unordered)
[65] Dorsal vertebrae with connective tissue bridging transverse processes: absent (0), present (1)
[66] Lateral process of coracoid: absent (0), present (1)
[67] Toes (0) shorter, (1) longer than metatarsus

5

10

15

20

25

30

35

40

45

50

55

60

65

67

allzero outgroup

00000

00000

00000

00000

00000

00000

00000

00000

00000

00000

00000

00000

00100

00

Scansoriopterygidae

??0?0

?????

00?00

000?0

???10

0???0

0?000

00010

00??{03}

10?0?

100??

0?000

??020

00

Archaeopteryx

00000

00?10

00000

00000

00000

00000

00000

0010?

00003

10000

1010?

(01)?000

11010

01

Rahonavis ostromi

?01?2

???1?

?????

?????

00???

0??00

000?0

01111

?0000

1??00

1010?

??00?

?1010

?1

Shenzhouraptor sinensis

1?1?{123}

0001?

00010

00??0

??001

0??00

0?000

011{01}?

00000

10000

101??

10001

?1010

01
Jixiangornis orientalis 0???? ????? ?0100 2???0 ??000 ????0 0???? 0211? 00??{01} 10100 ?0??? 0???? 1?010 01

Hulsanpes perlei

?????

?????

?????

?????

?????

????0

00???

0???1

?0?01

?????

?????

??0??

?????

??

Yandangornis longicaudus

1?1??

?????

??0?{01}

????0

???0?

00?00

0??00

0?110

00001

??1?{01}

00??0

??00?

?????

?0

Sapeornis chaoyangensis

0?000

???00

00?02

01000

00101

0?000

0??00

02000

00000

00?01

11(01)0?

0?0?1

?1200

01
Omnivoropteryx sinousaorum 0?0?? ????? ????1? ????0 ???00 0???0 0???? 1?10? ?0??? ????1 ????? ??0?1 ????0 ?1

Confuciusornis sanctus

10100

00?00

00(01)00

10001

00200

0?000

00000

12110

00001

00101

01010

10001

10200

01

Jibeinia luanhera

0?10?

???{12}?

??011

2?1??

??201

0????

0??1?

031{01}0

00??1

00101

11??1

1?0??

??{12}0?

?1

Protopteryx fengningensis

01101

?0???

?1102

21101

?0210

0???1

0000?

0201?

00001

00111

11???

1?0?1

??200

0?

Longipteryx chaoyangensis

0?102

?0???

01102

2?1?1

??211

0????

0??1?

03???

0000{01}

00101

?10?1

1?0?1

1?{12}?0

0?

Eocathayornis walkeri

01?02

?11{12}0

??112

21111

?1{12}11

?????

?????

?????

0????

?010?

????1

1???1

????0

0?

Iberomesornis romerali

?010?

?????

?11??

20001

??2??

?1??1

0??1?

03100

?00?1

0?10?

?10??

1?0??

{12}0200

00

Noguerornis gonzalezi

???0?

?????

11???

?10?1

?0?1?

?????

?????

?????

0????

?0???

?????

?????

?????

??

Halimornis thompsoni

?1??2

???21

?????

?111?

??2??

??0??

?????

?????

?????

0????

?????

?????

??{01}00

??

Cuspirostrisornis houi

0?1??

???{12}?

??22?

2?01?

??21?

0????

1??1?

?{234}?2?

00?0?

001?1

0????

1?0??

??{12}??

01

Concornis lacustris

?1102

10?2?

1112?

21111

1???1

1?1?1

1?011

0??20

000?1

?0101

0??11

1?0?1

?????

00

Sinornis santensis

01102

10010

?1123

21111

?1211

01011

1101?

0310?

00001

00101

01011

1?0?1

?0200

01
Longirostravis hani 0?1?? ???1? 111?? 21??? ??11? ???10 0???? 12??? 00??1 001?? 010?? 1?0?? ??100 0?
Vescornis hebeiensis 0???{123} ????1 1112{23} 21111 ??211 ??1?1 0?1?0 13010 ?1??1 0010? 1???1 1?001 ??20? 01

Gobipteryx minuta

1?1?2

?001?

11???

2????

?111?

1??01

010?0

1{45}???

?0?0?

0??01

?????

100??

?????

0?

Lectavis bretincola

??1??

?????

?????

?????

?????

???0?

1???0

?????

?{01}?11

?????

?????

??0??

?????

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Soroavisaurus australis

??1??

?????

?????

?????

?????

???11

111?1

1??2?

?0?01

?????

?????

??0??

?????

??

Yungavolucris brevipedalis

?????

?????

?????

?????

?????

????1

10??0

0???1

?0?01

?????

?????

??0??

?????

??

Avisaurus gloriae

?????

?????

?????

?????

?????

????1

11??1

1????

?1?01

?????

?????

??0??

?????

??

Enantiornis leali

???02

10021

?????

21110

11?1?

?????

?????

?????

00???

?0?0?

???01

1????

?????

0?

Neuquenornis volans

?1??2

10111

102??

2???1

?1?1?

?11?1

111?1

0??2?

000??

?0101

????1

1??01

?????

0?

Boluochia zhengi

1?1??

?????

??1??

?????

??2??

0??11

00?1?

0????

?0??1

001??

0????

??0??

??20?

?1

Otogornis genghisi

???0{34}

1????

???2?

211?1

???0?

?????

?????

?????

0????

???0?

????1

1????

?????

0?

Alexornis antecedens

????2

???{12}?

?????

2????

1????

?????

?????

?????

??0??

???0?

?????

1????

?????

??

Eoalulavis hoyasi

???02

1011?

11123

21111

?1?11

?1???

?????

?????

0????

?010?

??001

1???1

2???0

0?

Eoenantiornis buhleri

0?10?

?0???

11?1{23}

{12}????

?1211

01??1

0??1?

?{123}???

00??1

0010?

?1???

1???1

???0?

0?

unnamed Lebanon

?????

10?20

?1???

2????

?????

????1

0????

0??2?

?0??1

??10?

?????

1?0??

?????

01

Liaoningornis longidigitis

??1??

?????

??2??

?????

?????

0???0

0??1?

1?00?

?1??1

??1??

0????

??0??

?????

01

Apsaravis ukhaana

?0113

101{12}?

??2??

20111

003??

1??10

0???0

150?0

?2112

01102

01011

1?0?1

2?{12}{01}?

0?

Ambiortus dementjevi

???1{34}

0????

002??

200??

????1

?????

?????

?????

?????

??11?

?????

1???1

?????

??

Yanornis martini

0?1?{34}

?0???

?0203

2?0?0

??301

1???0

0?0?0

0410?

12??2

00111

?????

1???1

2??{01}?

01

Yixianornis grabaui

0?1?{34}

?0???

?0213

20??0

??301

1???0

0?0?0

0410?

12?02

01111

??0??

1?0?1

???{01}?

01

Patagopteryx deferrariisi

?0103

00?21

?????

200?1

00?00

1?010

000?0

04110

12011

0??02

0?01?

10?00

20{12}?0

?1

Vorona berivotrensis

??1??

?????

?????

?????

?????

?0110

00?00

0???0

?2001

?????

0????

??00?

?????

??

Songlingornis linghensis

0???{34}

01???

002??

2????

?????

?????

?????

?????

?{12}???

??11?

?????

1????

?????

0?

Gansus yumenensis

??1??

?????

?????

?????

?????

???00

00??0

0?110

?2?12

?????

?????

??10?

??????

?1

Hesperornithes

101?3

01?0?

000?3

10000

??30?

10000

00010

05100

?2112

0?112

010?0

0(01)11?

20200

01

Ichthyornis dispar

10114

0100?

00223

20010

00301

10000

00010

15???

12112

01112

01011

11111

20201

1?

Anas platyrhynchos

10114

01?1?

00223

20000

?0301

10000

00010

05000

12112

01112

01011

11111

20201

11

Microraptor

00000

???1?

00000

0?0?0

??000

00?00

0??00

00111

00003

10(01)00

100?1

??0?1

11020

00

Caudipteryx

0?000

???0?

00002

0???0

??{01}00

0??00

0?000

00?00

00003

00000

000??

0?0?0

?0000

00

basal Troodontidae

00000

???{12}?

0?000

000?0

?0?00

00000

001?0

10111

00003

10?00

?000?

??000

1(01)000

00

{} = uncertainty (one of the character states occurs)
() = polymorphism (all character states occur)

Taxon comments:

Scansoriopterygidae: Composite of Scansoriopteryx heilmanni and Epidendrosaurus ningchengensis – very similar, identical apart from "?" in this matrix.
Archaeopteryx: A. lithographica and A. bavarica together. Only difference except for position of "?" : A. lithographica has 56(0), A. bavarica has (1). Therefore polymorphism.
Iberomesornis romerali: Character 67 after measurements by Sereno 2000. Consistent with longish mt; impossible to see where bone ends and horn sheath continues alone in illustrations.
Hesperornithes: Composite of Hesperornis regalis and Baptornis advenus. Differences in missing data and inapplicable codings, and perhaps in character 57.
Microraptor: Composite of Microraptor zhaoianus and Microraptor gui – identical apart from "?" in this matrix.
Caudipteryx: Species-level taxonomy (C. zoui, C. dongi, C. sp.) uncertain and ignored. All differences would be in the positions of "?" .
basal Troodontidae: Coded after Sinovenator and Sinornithoides. Only difference except for position of "?" : -venator has 62(1), -ornithoides appears to have (0). Therefore coded as polymorphism.

Character comments:

8: In case of 65(0) the theoretical position of the coracoid fossa was guessed.
13: In absence of ossified sternum coded as 0.
52: If not irreversible, Yandangornis likes to cluster with Confuciusornis. That said, it likes to cluster with C. anyway.
53: Preacetabular process measured from the cranialmost point of the ilium to the cranial border of the acetabulum. Postacetabular process measured from the caudalmost point of the ilium to the caudal border of the ischiadic peduncle of the ilium.
61: In the absence of visible ilia, coded from the length of the sacral ribs.
63: Ordered in this way – plesiomorphy = (1) – to spare me a stepmatrix. Occasionally the transition point is in the pygostyle.
64: Should perhaps be ordered like 63... Pygostyles coded as (0) because of Halimornis, Sinornis, Boluochia? etc..

Stepmatrices:

character 5     character 23     character 45
0 1 2 3 4         0 1 2 3          0 1 2 3
- 1 1 2 3         - 1 2 2          - 1 2 1
1 - 1 1 2         1 - 1 1          1 - 1 2
1 1 - 2 3         2 1 - 1          2 1 - 1
2 1 2 - 1         1 1 2 -          1 2 2 -
3 2 3 1 -

Cladograms:

Yeah, er. Coming.

Strict consensus when all taxa are included and when Alexornis, Hulsanpes, Lectavis and Yungavolucris are deleted: Grass.


Spambot, terminate yourself.

Questions? Including requests for the NEXUS file?

I said: Spambot, terminate yourself.


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